'Orsten' Taxa (List Version, preliminary)
List of 'Orsten' Taxa (from Australia, China, Poland, Sweden, and UK so far)
This is our first, almost complete list of more than 50 'Orsten' fossil taxa (name giving also by others, of course) and the only authorized by us. Current update from April 15, 2011
Note that Agnostos pisiformis is different from the rest of 'Orsten' arthropods in so far that it has been known for long in form of its calcitic head and tail shields of the large specimens. What is still missing in this list are the cambrogeorginids, palaeoscolecids and more non-arthropods and small shellies, which will be added in due course.
We list the following items:
- ▪ Number
- ▪ Taxon (mainly species)
- ▪ Locality/stratigraphic range
- ▪ Systematics
- ▪ Major reference(s)
- ▪ Picture if available
You can also download a table of 'Orsten Taxa in PDF format. It will display in a new browser window.
In this Section...
1) Aengapentastomum andresi Waloszek, Repetksi & Maas, 2006
- ▪ Västergötland, Sweden, Furongian, uppermost Cambrian, reworked into Lower Ordovician layers
- ▪ Arthropoda stemlineage, Pentastomida stemlineage
- ▪ Waloszek et al. 2006
2) Agnostus pisiformis (Wahlenberg, 1818)
- ▪ Skåne, Öland, Väster-götland, Sweden, Cambrian series 3, uppermost Middle Cambrian, according to latest decisions of the International Stratigraphic Commission, A.-pisiformiszone (originally lowermost stratum of Upper Cambrian of the Baltoscandic Region)
- ▪ Arthropoda s. str. Euarthro-poda incertae sedis, possibly closer to Crustacea s. l. (sister taxon?)
- ▪ Müller & Walossek 1987, Walossek & Müller 1990, 1998a, Stein et al. 2005
3) Boeckelericambria pelturae Walossek & Müller, 1994
- ▪ Kinnekulle, Västergötland, Sweden, Furongian = uppermost Cambrian, according to Internl. Strat. Comm., zone 5/6 of original Upper Cambrian of Baltoscandic Region
- ▪ Arthropoda stemlineage, Pentastomida stemlineage
- ▪ Walossek & Müller 1994
4) ???
5) Bredocaris admirabilis Müller, 1983
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda, Thecostraca stemlineage
- ▪ Müller 1983, Müller & Walossek 1988b, Walossek & Müller 1992, 1998a, b
6) Cambrocaris baltica Walossek & Szaniawski, 1991
- ▪ Furongian, late Cambrian of northern Poland
- ▪ Euarthropoda Crustacea stemlineage
- ▪ Walossek & Szaniawski 1991
7) Cambropachycope clarksoni Walossek & Müller, 1990
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Middle Cambrian
- ▪ Euarthropoda, Crustacea stemlineage
- ▪ Walossek & Müller 1990
8) Cambropycnogon klausmuelleri Waloszek & Dunlop, 2002
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3, uppermost Middle Cambrian
- ▪ Euarthropoda Chelicerata s.str. Pycnogonida stemlineage
- ▪ Müller & Walossek 1986b, 1988a, Walossek & Dunlop 2002, Vilpoux & Waloszek 2003
9) Dala peilertae Müller, 1983
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian, late Cambrian
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda stemlineage
- ▪ Müller 1981, see also Müller 1983, which was not the real first description
10) Falites fala Müller, 1964, ST 1708 (UB W 237)
- ▪ Sweden, Ctenopyge Zone (Zone 5), Västergötland
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Müller 1964, see also 1979 and Maas et al. 2003
11) Goticaris longispinosa Walossek & Müller, 1990
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian, late Cambrian Euarthropoda
- ▪ Crustacea stemlineage
- ▪ Walossek & Müller 1990
12) Haffnericambria trolmeniensis Walossek & Müller, 1994
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian, late Cambrian
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek & Müller 1994
13) Henningsmoenicaris scutula (Walossek & Müller, 1990) new name for Henningsmoenia scutula Walossek & Müller 1990
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3, uppermost Middle Cambrian Euarthropoda
- ▪ Crustacea stemlineage – Hennings-moenia is pre-occupied by Schallreuter (1968)
- ▪ Walossek & Müller, 1990, 1991
14) Hesslandona angustata Maas, Waloszek & Müller, 2003, ST 3122 (UB W 203, Holotype)
- ▪ A. pisiformis Zone, Gum, Västergötland, S Euarthropoda
- ▪ Crustacea Labrophora Phosphatocopina
- ▪ see Maas et al. 2003
15) Hesslandona curvispina Maas, Waloszek & Müller, 2003, ST 3721 (UB W 205, Holotype)
- ▪ A. pisiformis Zone, Kinnekulle, Västergötland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Maas et al. 2003
16) Hesslandona kinnekullensis Müller, 1964, ST 2225 (UB W 174)
- ▪ Olenus Zone (Lowermost Furongian) of Västergötland, Öland, S Euarthropoda
- ▪ Crustacea Labrophora Phosphatocopina
- ▪ Müller 1964, see also Maas et al. 2003
17) Hesslandona necopina Müller, 1964 (Type species of Hesslandona Müller, 1964), ST 2204 (UB W 165)
- ▪ A. pisiformis Zone, Västergötland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Müller 1964, see also Maas et al. 2003
18) Hesslandona suecica Maas, Waloszek & Müller, 2003, ST 3211 (UB W 197)
- ▪ A. pisiformis Zone, Kinnekulle, Västergötland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Maas et al. 2003
19) Hesslandona toreborgensis Maas, Waloszek & Müller, 2003, ST 3771 (UB W 214, Holotype)
- ▪ Olenus Zone (Lowermost Furongian) of Västergötland, Öland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Maas et al. 2003
20) Hesslandona trituberculata Lochmann & Hu, 1960, ST 3244 (UB W 179)
- ▪ Olenus Zone (Lowermost Furongian) of Kinnekulle, Västergötland, Öland, S; Holotype from Du Noir Limestone, early Upper Cambrian, Cedaria Zone at Sheep Mountain Section, Northwest Wind River Mountains, northeastern Sublette County, Wyoming, USA
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Lochmann & Hu 1960, see also Maas et al. 2003
21) Hesslandona unisulcata Müller, 1982 ST xxxx
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3, uppermost Middle Cambrian
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Müller 1979, Müller 1982, Maas et al. 2003, Maas & Waloszek 2005
22) Hesslandona ventrospinata Gründel in Gründel & Buchholz, 1980, ST 3488 (UB W 186)
- ▪ A. pisiformis Zone, Västergötland, S; Holotype from erratic boulders of Rügen
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Gründel & Buchholz 1980, see also Maas et al. 2003
23) Heymonsicambria gossmannae Walossek & Müller, 1994
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian, late Cambrian
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek & Müller 1994
24) Heymonsicambria kinnekullensis Walossek & Müller, 1994
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek & Müller 1994
25) Heymonsicambria repetskii Walossek & Müller, 1994
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek & Müller 1994
26) Heymonsicambria scandica Walossek & Müller, 1994
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek & Müller 1994
27) Heymonsicambria sp. a of Walossek & Müller (1994)
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Furongian
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek & Müller 1994
28) Heymonsicambria taylori Walossek, Repetski & Müller, 1994
- ▪ Upper Furongian or Lower Ordovician, Newfoundland, Canada
- ▪ Arthropoda stemlineage Pentastomida stemlineage
- ▪ Walossek, Repetski & Müller 1994
29) Klausmuelleria salopensis Siveter, Waloszek & Williams, 2003
- ▪ Comley, UK, possibly series 2, uppermost Lower Cambrian Arthropoda
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina – sister taxon to all others
- ▪ Siveter et al. 2001, 2003, Maas et al. 2003, Maas & Waloszek 2005
30) Markuelia lauriei Haug, Maas, Waloszek, Donoghue & Bengtson, 2009
- ▪ Rogers Ridge, Australia, possibly, series 2 or 3, former Middle Cambrian
- ▪ Nemathelminthes ?Cycloneuralia
- ▪ Haug et al. 2009
31) Martinssonia elongata Müller & Walossek, 1986
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea stemlineage
- ▪ Müller & Walossek 1986, Walossek & Müller 1990
32) Musacaris gerdgeyeri Haug, Waloszek & Maas, 2010
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea stemlineage
- ▪ Haug et al. 2010a
33) Oelandocaris oelandica Müller, 1983
- ▪ Öland, Sweden, uppermost Furongian, new material from Västergötland, which is also uppermost Cambrian
- ▪ Euarthropoda Crustacea stemlineage
- ▪ Müller 1983, Stein et al. 2005
34) Orstenocarida Müller & Walossek, 1988
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Cambrian = Furongian
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda Thecostraca stemlineage
- ▪ see Bredocaris Müller 1983, Müller & Walossek 1988b, Walossek & Müller 1992, 1998a, b
35) Orstenoloricus shergoldii Maas, Waloszek, Haug & Müller, 2009
- ▪ Rogers Ridge, Australia, possibly, series 2 or 3, former Middle Cambrian
- ▪ Nemathelminthes Cycloneuralia Scalidophora
- ▪ Maas et al. 2009
36) Orstenotubulus ewamuellerae Maas, Mayer, Kristensen & Waloszek, 2007
- ▪ Skåne, Oeland, Väster-götland, Sweden, Cambrian series 3
- ▪ Arthropoda stemlineage
- ▪ Maas et al. 2007
37) Phosphatocopina Müller, 1964
- ▪ worldwide, Middle to Upper Cambrian, according to the find of Klausmuelleria also Lower Cambrian
- ▪ Arthropoda Euarthropoda Crustacea Labrophora – sister taxon to Eucrustacea
- ▪ see Hesslandona Müller 1964, 1979a, b, c, 1982, Maas et al. 2003, 2004, Maas & Waloszek 2005
38) Rehbachiella kinnekullensis Müller, 1983
- ▪ Kinnekulle, Västergötland, Sweden, uppermost Middle Cambrian according to latest decisions of the International Stratigraphic Commission, Agnostus-pisiformis zone (originally lowermost stratum of UC of the Baltoscandic Region)
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Branchiopoda – sister taxon to Eubranchiopoda
- ▪ Müller 1983, Walossek & Müller 1990, 1998a, b, Walossek 1993, 1995, 1996, 1999, Walossek 2003a, b, Waloszek & Maas 2005
39) Sandtorpia vestro-gothiensis Haug, Maas & Waloszek, 2010
- ▪ Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea stemlineage
- ▪ Haug et al. 2010b
40) Shergoldana australiensis Maas, Waloszek, Haug & Müller, 2007
- ▪ Rogers Ridge, Australia, possibly, series 2 or 3, former Middle Cambrian
- ▪ Nemathelminthes ?Cycloneuralia
- ▪ Maas et al. 2007
41) Skaracarida Müller & Walossek, 1985
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda copepodan lineage (related to Mystacocarida and Copepoda) see Skara
- ▪ Müller & Walossek 1985
42) Skara anulata Müller 1983
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda copepodan lineage Skaracarida
- ▪ Müller & Walossek, 1985 Müller 1983, Walossek & Müller 1990, 1998a, b,
43) Skara ?minuta
- ▪ from a borehole in Poland, straum unclear
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda copepodan lineage Skaracarida
- ▪ Müller & Walossek, 1985 Walossek & Szaniawski 1991
44) Skara minuta Müller & Walossek, 1985
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda copepodan lineage Skaracarida
- ▪ Müller & Walossek, 1985 Müller & Walossek 1985, Walossek & Müller 1990, 1998a, b
45) Skara huanensis Dong in Liu & Dong, 2007
- ▪ China, Upper Cambrian = Furngian, but unclear
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca Maxillopoda copepodan lineage Skaracarida
- ▪ Müller & Walossek, 1985 Liu & Dong 2007
46) Trapezilites minimus (Kummerow, 1931), ST 3423 (UB W 220)
- ▪ A. pisiformis Zone and Olenus Zone (Lowermost Furongian) of Västergötland, Öland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
47) Type A larva, no 1
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3, but few specimens also from uppermost Furongian, more lower Ordovician (Canada) and series 2/3 (Australia)
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea
- ▪ Müller & Walossek 1986b
48) Type A larva, no 2
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea
- ▪ Walossek & Müller 1989
49) Veldotron bratteforsa Müller, 1964, ST 3491 (UB W 233)
- ▪ Olenus Zone (Lowermost Furongian) of Västergötland, Öland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Müller 1964, see also Maas et al. 2003
50) Vestrogothia spinata Müller, 1964, ST 2252 (UB W 243)
- ▪ Ctenopyge Zone (Zone 5), Västergötland, S
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Müller 1964, see also Maas et al. 2003
51) Waldoria rotundata Gründel in Gründel & Buchholz, 1981, ST 11091 (UB W 224)
- ▪ A. pisiformis Zone, Västergötland, S; Holotype from erratic boulders of Rügen
- ▪ Euarthropoda Crustacea Labrophora Phosphatocopina
- ▪ Gründel & Buchholz 1980, see also Maas et al. 2003
52) Walossekia quinquespinosa Müller, 1983
- ▪ Kinnekulle, Västergötland, Sweden, Cambrian series 3
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca incertae sedis
- ▪ Müller 1983
53) Wujicaris muelleri, Zhang, Maas, Haug, Siveter & Waloszek, 2010
- ▪ Xiao-tan section, Yongshan, Yunnan Province, China (Yu’anshan Formation, Eoredlichia-Wutingaspis Biozone, upper Lower Cambrian of traditional usage)
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca incertae sedis
- ▪ Zhang et al. 2010
54) Yicaris dianensisZhang, Siveter, Waloszek & Maas, 2007
- ▪ Province Yunnan, China, Cambrian series 1
- ▪ Euarthropoda Crustacea Labrophora Eucrustacea Entomostraca incertae sedis
- ▪ Zhang, Siveter, Waloszek & Maas, 2007
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